Plants

Since New Zealand became isolated from Gondwanaland approximately 80 - 90 million years ago, it has many archaic plants and animals, all of which have evolved in the absence of mammals. Many of New Zealand's forests are unique, such as the kauri and podocarp forests which date back 200 million years. Below we present a brief overview of New Zealand's prominent native vegetative cover.
 
 

Kauri Forest | Podocarp Forest | Beech Forest | Mountain Cedar | Alpine Plants

 

 

Mangrove

The mangrove (Avicennia marina spp. australasica) is distributed around the top half of the North Island, barely reaching past the winter isotherm of 0°C. This is south to Kawhia harbour on the west coast, and Opotiki in the Bay of Plenty. From Whangarei northwards, trees can reach a height of 9 metres, especially on the edges of tidal courses where there is little wave action. However, on large tidal flats, the mangrove generally grows as a small bush, no more than half a metre in height. It does best in areas where there is protection from wave action, such as behind shell banks. They grow best in areas where the water depth is approximately half a metre at high tide. In general, they are the first pioneering plant to stabilise mudbanks at river entrances and to colonise harbours.

The leaves of the mangrove are typical of xerophytes, they are sclerified with a shiny upper surface and white tomentum underneath to reflect the intense light coming off the water. Its root system is very strong, and forms a horizontal network, just under the mud surface. The vertical pencil-like aerial breathing roots, ‘pneumatophores’ stick out of the mud for some 10 cm. These have a water-resistant bark and a spongy aerenchyma, stiffened by an axis of vascular tissue. These pneumatophores are often coated with the estuarine barnacle, and black mussel. Rock oysters can also encrust the pneumatophores and trunk of the mangrove, so much so, that their weight can bend the pneumatophore over. This sheltered area of water underneath the mangrove canopy also provides an important and protective breeding ground for crustaceans and fishes.
 
 
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Forests

The forests of New Zealand are evergreen rainforests. There are three major forest classes: the lowland species-rich kauri forest north of c. 38° S; mixed broadleaf-podocarp forest (angiosperm-conifer) that dominates the lowlands south of c. 38° S; species-poor beech, and mountain cedar occupy much of the montane zone.
 
 

Lowland Kauri forest north of c. 38° S

The complex structure of the kauri (Agathis australis) forest (including nikau palms) gives it a subtropical appearance. In this forest type, kauri forms an emergent layer above a mixed angiosperm canopy. Vegetative growth is significantly reduced underneath the spreading canopy of the mature kauri, due to low light levels, and the kauri’s production of slowly decomposing leaf litter that releases few nutrients, leading to the formation of mor humus and soil podzolisation. Dense stands of kauri, with multiple-aged structure are most commonly found on ridges. This is due to such sites suffering more frequent environmental disturbance through windfall and soil slipping, allowing light-demanding kauri seedlings to become established. Kauri are more adapted to the infertile and more drought-prone soils of ridge sites, whereas in gullies (fertile sites) the angiosperm forest forms a continuous canopy matrix that excludes kauri.
 
 

Lowland Podocarp hardwood forests south of c. 38° S

These lowland forests occupy c. 500 000 ha in the North Island and 400 000 ha in the South Island. The largest trees are typically emergent podocarps, rimu, kahikatea, miro, matai, totara densely stocked on recent pumice, alluvial and glacial soils, and emergent but more sparsely stocked on older, usually hill-country sites.Associated angiosperms (‘hardwood’ or ‘broad-leaf’ trees) are commonly tawa, ratas, kamahi, rewarewa, kohekohe, whiteywood. Forest type and class structure varies according to species composition, altitude, topography, soil type, climate, and historical disturbance.
 
 
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Montane forests

Nothofagus (Beech Forest)

In New Zealand, the montane forest is predominantly Nothofagus "southern beech". Five taxa of beech are currently recognised in New Zealand. These comprise four species, one of which has two varieties. They are: red beech (N. fusca), hard beech (N. truncata), silver beech (N. menziesii), black beech (N. solandri var. solandri), and sub-species mountain beech (N. solandri var. cliffortioides). The beech can reach a reasonable size, the largest being red beech recorded up to 42.7 m in height, with a trunk 3.0 m in diameter. Growth form varies considerably between species, altitude and site conditions.

The beech species are adapted to growing on low fertility and poorly drained soils, dry sites such as ridge crests, and areas of recent disturbance. They often form extensive, relatively uniform forests in the montane region, but at lower altitudes give way to the more competitive hardwoods and softwoods. The beech species tend to form discrete pure stands rather than occur in intricate mixtures with other tree genera. This may be related to mycorrhizal symbionts.

Beech forest occurs almost throughout the full latitudinal range of the North and South Islands, though its distribution in the north-west tends to be local. It does not occur naturally on the neighbouring Stewart Island, nor does it occur on the subantarctic islands south of New Zealand such as the Aucklands, Campbell and Snares, even though these fall well within the latitudinal range of the genus in South America.

The altitude of the beech timberline varies depending on latitude, from 600-1400 m a.s.l.. Hard beech has a more northerly distribution and is associated with lower altitudes than the other beeches, whereas mountain beech has the highest altitudinal range of any beech species. At lower altitudes, beech is associated with the podocarp species, particularly rimu, and the broad-leaved hardwoods, kamahi, and in the North Island, tawa. Within their altitudinal range, beech generally form pure forest stands. In montane and subalpine regions on shallow and poorly drained soils, beech is sometimes associated with cedar and shrub Dacrydium species. At their upper altitudinal range, on exposed ridges, beech tend to become dwarfed with wind-shorn canopies. On the most exposed sites, silver beech can be reduced to flattened cushions with slender layering branches, sometimes less than 50-100 cm high and 15 cm diameter. In such sites, beech can develop specialised growth forms as a result of drifting and avalanching snow.
 
 

Mountain cedar

The mountain cedar (Libocedrus bidwillii) with associated podocarps and some anigosperms make up the subalpine forests where Nothofagus spp. are absent. The climate of this forest is typified by high rainfall, frequent fogs, short cool summers and wet organic soils. Stands of cedar predominantly consist of ancient gnarled individuals (often older than 700 years plus), with relatively few younger trees, saplings or seedlings. This even age distribution is due to spasmodic regeneration following canopy disturbance after windthrow or mass movement on steep slopes. The age distributions of some stands can reflect the effects of disturbances that occurred about 200-400 years ago.
 
 
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Alpine Plants

The mountains supporting alpine plants can be found on Te Moehau and the Raukumara Range in the north, through to Stewart Island in the South. The evolution of these mountains has varied considerably over the last 20 million years. Their parent material, past and present climates and degree of isolation has had a major influence on alpine communities. The present relief patterns of these mountains has been largely a result of erosion by water; in alpine regions, wind and ice with freeze-thaw action are responsible. Parent rock material varies between regions of uplift and its degree of erodibility affects vegetation. For example, Greywacke (metamorphosed sedimentary rock) is particularly prone to frost shattering and is eroded rapidly, resulting in the development of scree slopes.

The North Island volcanic mountains supporting alpine communities are: Egmont 2,518 m (8,260 ft) on the west coast; Ruapehu, 2,796 m (9,175 ft), Tongariro 1,986 m (6,517 ft), and Ngauruhoe 2,290 m (7,515 ft) on the central plateau. These volcanoes reached their present height during the last glaciation in the Pleistocene, except Ngauruhoe which is younger. Their relatively early evolution has meant that alpine community diversity is relatively low compared with other alpine areas. The highly erodible porous larva and loose scoria makes it difficult for plants to become established.

About 93% of the alpine plant species are endemic to New Zealand; these are considered to have originated either here or on the Antarctic continent. New Zealand’s alpine flora has probably evolved within the last two million years or so during the last glaciations, from pre-existing nonalpine flora, together with chance immigrants from other areas that supported alpine plants, such as Antarctica, or Australia. This theory of the recent evolution of alpine flora, is supported by the widespread occurrence of hybridism and the variation or polymorphism in many of the alpine plant groups.

In the 1000 m altitudinal range between the tree line and snow line, there is a greater range of alpine vegetation than occurs in most other parts of the world. Because of the great plant diversity in this area, two zones are recognised. The low-alpine zone is dominated by tussocks, Chionochloa spp., Coprosma, Dracophyllum, Hebe, Celmisia, Ranunculus, Gentiana, Aciphylla, Anisotome, and Astelia. Some of these are represented in the high-alpine zone where there is less complete plant cover. The transition between the low and high-alpine zones is generally too subtle to detect, however, it mostly occurs half way between the tree and snow lines. Plants in the high-alpine zone tend to grow in a compact creeping form (cushion) close to the ground, such as the Raoulia (vegetable sheep). The ground in this zone can be very mobile in areas of scree, sloping 30 - 35 degrees. However, it is generally only the top 10 - 20 cm layer that is mobile. Beneath is a compact, permanently moist soil, where highly adapted plants with large taproots or rhizomes can persist. The leaves these plants do not persist over winter, and are delicately attached to the stem so that they break off easily without damaging.

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